Definition of grana in English:

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Pronunciation: /ˈɡrɑːnə/

plural noun (singular granum)

The stacks of thylakoids embedded in the stroma of a chloroplast.
Example sentences
  • Chloroplast envelopes, stacked grana and unstacked stroma thylakoids were well-defined.
  • The chloroplasts with large grana stacks were seen in 50% of cases but none of the parameters significantly explained the occurrence.
  • The VPC chloroplasts possessed well-developed thylakoids with grana.


Late 19th century: plural of Latin granum 'grain'.

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